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Selaginella sp.식물/들꽃-부처손과(Selaginellaceae) 2023. 6. 19. 16:38
국표에는 Selaginella (부처손속)이 28종 이다.
Selaginella is the sole genus in the family Selaginellaceae, the spikemosses or lesser clubmosses, a kind of vascular plant.
This family is distinguished from Lycopodiaceae (the clubmosses) by having scale-leaves bearing a ligule and by having spores of two types. They are sometimes included in an informal paraphyletic group called the "fern allies". S. moellendorffii is an important model organism. Its genome has been sequenced by the United States Department of Energy's Joint Genome Institute. The name Selaginella was erected by Palisot de Beauvois solely for the species Selaginella selaginoides, which turns out (with the closely related Selaginella deflexa) to be a clade that is sister to all other Selaginellas, so any definitive subdivision of the species into separate genera leaves two taxa in Selaginella, with the hundreds of other species in new or resurrected genera.
Selaginella occurs mostly in the tropical regions of the world, with a handful of species to be found in the arctic-alpine zones of both hemispheres. Fossils assignable to the modern genus are known spanning over 300 million years from the Late Carboniferous to the present.
Description
Selaginella species are creeping or ascendant plants with simple, scale-like leaves (microphylls) on branching stems from which roots also arise. The stems are aerial, horizontally creeping on the substratum (as in Selaginella kraussiana), sub-erect (Selaginella trachyphylla) or erect (as in Selaginella erythropus). The vascular steles are polystelic protosteles. Stem section shows the presence of more than two protosteles. Each stele is made up of diarch (having two strands of xylem) and exarch (growing outward in) xylems. The steles are connected with the cortex by means of many tube-like structures called trabeculae, which are modified endodermal cells with casparian strips on their lateral walls. The stems contain no pith.
In Selaginella, each microphyll and sporophyll has a small scale-like outgrowth called a ligule at the base of the upper surface. The plants are heterosporous with spores of two different size classes, known as megaspores and microspores.
Unusual for the lycopods, which nearly always have microphylls with a single unbranched vein, the microphylls of a few Selaginella species contain a branched vascular trace.
Under dry conditions, some species of Selaginella can survive dehydration. In this state, they may roll up into brown balls and be uprooted, but can rehydrate under moist conditions, become green again and resume growth. This phenomenon is known as poikilohydry, and poikilohydric plants such as Selaginella bryopteris are sometimes referred to as resurrection plants.
Species
Main article: List of Selaginella speciesThere are about 750 known species of Selaginella. They show a wide range of characters; the genus is overdue for a revision which might include subdivision into several genera. Better-known[by whom?] spikemosses include:
- Selaginella apoda – meadow spikemoss; eastern North America
- Selaginella arizonica Maxon – west Texas to Arizona and Sonora, Mexico
- Selaginella asprella
- Selaginella bifida – Rodrigues Island
- Selaginella biformis
- Selaginella bigelovii
이하 생략
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