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Nepenthes hamata J.R.Turnbull & A.T.Middleton식물/들꽃-벌레잡이풀과(Nepenthaceae) 2023. 5. 6. 12:20
국표에 없다.
Nepenthes hamata /nɪˈpɛnθiːz həˈmɑːtə/ is a tropical pitcher plant endemic to Sulawesi, where it grows at elevations of 1400–2500 m above sea level.
The specific epithet hamata is derived from the Latin word hamatus, meaning "hooked". It describes the appearance of the highly developed peristome teeth of this species.
Nepenthes hamata is a strong climber. The stem, which may be branched, reaches a maximum length of around 7 m. It is terete to obtusely trigonous and varies in diameter from up to 3 mm in rosettes and short stems, to 4–5 mm in climbing stems. In the former, the internodal length is typically up to 6 mm and in the latter 3.5–6 cm.
Leaves
Leaves are sessile and chartaceous in texture. The shape of the lamina (leaf blade) is variable: it may be linear, lanceolate, or slightly elliptic.
In the case of rosettes and short stems, the lamina is typically oblanceolate to oblong-elliptic and measures up to 7.5 cm in length by 2.5 cm in width. It has an acute apex and does not exhibit a peltate tendril attachment. The laminar base is amplexicaul, clasping the stem and giving it a subperfoliate appearance. Auricles may be present, although their level of development varies. The lamina may be slightly decurrent down the stem, but not prominently so.
Laminae produced on climbing stems are predominantly oblong-elliptic in shape, but may rarely be lanceolate. They are larger than those borne on shorter stems, but nonetheless relatively short, measuring 5–15 cm in length by 1.8–4 cm in width. The laminar apex may be acute to obtuse, while the base is abruptly contracted and clasps the stem; it may be decurrent for up to 1.5 cm in some populations. Two (rarely up to four) longitudinal veins are present on either side of the midrib. They are restricted to the outer third to half (rarely up to two-thirds) of the laminar surface. Pinnate veins are inconspicuous; they are patent and branching.
Pitchers
Rosette and lower pitchers are only produced for a short time before the plant transitions into a scrambling vine (although they may be produced later on offshoots from the original stem). They grow up to 18 cm high by 5 wide. They are ovate in the basal fifth to half of the pitcher cup, being bulbous to varying degrees in this portion, often narrowly so. The pitcher is narrower in the upper part and may be cylindrical or slightly infundibular towards the orifice. The boundary between these two portions is often delineated by a pronounced hip. The ventral face of the pitcher is flattened. A pair of wings up to 16 mm wide runs down the ventral surface of the pitcher cup. These wings bear densely packed filiform fringe elements up to 19 mm long, which commonly exceed the width of the wings themselves. These filaments are often arranged in pairs, spaced around 2 mm apart, and are usually branched dichotomously once or twice. The waxy zone of the inner surface is well developed.
The pitcher mouth is often distinctly rhomboid in shape, but may also be oval or elliptic. It has a highly oblique insertion and is concave when viewed in profile. The mouth rises at the rear to form a tapered neck, which may be held upright or be inclined forward over the pitcher orifice. The species is noted for having possibly the most developed peristome in the genus. It is cylindrical to slightly flattened, glossy, rigid, and up to 5 mm wide (excluding the teeth). It consists of expanded ribs up to 6 mm high and spaced up to 5 mm apart. The entire flanges can measure up to 15 mm in length, although their size and number differ between populations (a typical number being approximately 20 on each side of the orifice). These flanges form exceptionally long, incurved teeth at the inner edge of the pitcher orifice. The teeth are sickle-shaped (falcate) and extend approximately 7 mm into the interior, as measured from the inner edge of the peristome to the tooth apex. The outer edge of the peristome is entire, with the recurved flanges extending for around 2 mm past the rim. The teeth of the neck may assume a dagger-like shape and measure up to 10 mm by 2 mm. These uppermost teeth are often noticeably splayed forward.
The pitcher lid or operculum is elliptic to ovate and measures up to 6 cm in length by 5 cm in width. It is held roughly horizontally and often has upturned margins. The apex is rounded, whereas the base is occasionally cordate[4] or subcordate. No appendages are present on the lower surface, but multicellular filiform appendages may be found on the upper surface, though this feature is often unstable and may be absent altogether. These 'hairs' or 'tentacles' are up to 20 mm long and may be branched. They arise from the ends of the lid veins and may number as many as 45 on each side of the lid. They are mainly restricted to the outer margins of the lid, where they are around 3 rows deep. The lower surface of the lid bears sparsely distributed nectar glands in the form of shortly elliptic, bordered pits. These are very small and inconspicuous, measuring only 0.1–0.2 mm in diameter. A spur up to 9 mm long is inserted near the base of the lid. It may be simple or repeatedly branched to the point of being fasciculate. The herbarium material examined by Cheek and Jebb exhibited spurs that were basally 5-branched, with each branch being secondarily ramified.
Upper pitchers are similar in shape to their terrestrial counterparts, though usually more elongated, growing to 7–25 cm in height by 1.2–6 cm in width. The basal fifth to third of the trap is ovate, narrowing and becoming cylindrical to slightly infundibular above. As in lower pitchers, a conspicuous hip often marks the boundary between these two parts. In most cases, a pair of narrow ribs is present in place of the wings, although fully developed fringed wings are sometimes encountered. The peristome is often even more developed in aerial traps, with longer and more widely spaced teeth, which curve completely outwards in some populations. These teeth are usually 12–16 mm long by 2–3 mm high, and spaced 2.5–6 mm apart. Upper pitchers resemble lower ones in most other respects.
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